The claims come amid questions over who initially paid for Johnson's apartment to be upgraded -- and they hit very close to home for the Prime Minister, directing public attention to both his former closest ally, Dominic Cummings, and his fiancée, Carrie Symonds.
Johnson, the outspoken leader whose early handling of the coronavirus pandemic divided Britons, is no stranger to backlash from across the political aisle.
But the latest episode marks the first time Cummings -- who left government late last year, months after a scandal over his movements damaged public trust in the UK's Covid-19 lockdown rules -- has leveled allegations, on the record, against Johnson's premiership.
Where does Boris Johnson live?
Compared with the Executive Residence at the White House or the various presidential palaces dotted around the world's capital cities, the British Prime Minister's London living quarters are relatively humble.
Leaders would traditionally live in a flat above Number 10 Downing Street, but recent Prime Ministers have instead moved into the larger apartment next door, at Number 11 -- above the offices of the Chancellor of the Exchequer.
Even so, the apartment is far from spacious. The buildings that line Downing Street consist of cramped offices and narrow corridors, and Prime Ministers are not afforded luxury in their official residence.
Prime Ministers are given £30,000 ($41,000) of public money a year to renovate the apartment during their term, and it is not uncommon for them or their spouses to put their own stamp on the interior design.
Tony Blair faced questions in 2002 over the cost of his revamp, which reportedly included adding a study. Pictures released during David Cameron's premiership showed a modern kitchen and open-plan living space, and Theresa May posed for a 2016 interview next to a fireplace, though it's not clear whether she made any significant changes to the interior design.
Of course, British leaders are not confined to this small apartment all year. The far more spacious country house residence of Chequers sits in Buckinghamshire, a short drive from London, and that destination is often used to entertain foreign leaders or to provide the Prime Minister with an escape from Westminster.
Why are Johnson's renovations controversial?
Johnson has been under pressure for weeks to explain how he paid for the renovations to his apartment, which British news outlets have reported cost around £200,000 ($280,000).
Those renovations were reportedly overseen by Johnson's fiancée, Carrie Symonds. But his former chief adviser, Dominic Cummings, who left his role late last year after amassing a reputation as the driving force behind the Prime Minister's Brexit policy and re-election, alleged in a blog post that Johnson planned to have Conservative Party donors pay for the upgrade.
If that was the case, the loan would need to have been declared to the UK's Electoral Commission. Political donations and loans are tightly controlled in the United Kingdom, with loans of more than £7,500 ($10,400) logged and publicly revealed by the commission four times a year.
"I told him I thought his plans to have donors secretly pay for the renovation were unethical, foolish, possibly illegal and almost certainly broke the rules on proper disclosure of political donations," Cummings wrote in an explosive post on Friday.
"I refused to help him organise these payments. My knowledge about them is therefore limited," Cummings also wrote.
In an interview with the BBC on Sunday, Johnson's international trade secretary, Liz Truss, said the Prime Minister had paid for the renovations himself. But she did not say whether he had borrowed the money from a donor first.
A Number 10 spokesperson told CNN that the government has "transparently laid out the historic expenditure" and that "any costs of wider refurbishment ... have been met by the Prime Minister personally."
The spokesperson added that more information on the works will be released when the Cabinet Office publishes its yearly accounts, which usually happens in July. However, a separate register of ministerial interests has not been published by the government since last July, despite the ministerial code stating that one should be made public "twice yearly."
These outstanding questions have combined to spark a political storm over Number 10, with critics and opposition MPs increasing calls for clarity over how the apartment was revamped.
What could happen now?
The opposition Labour Party has called for Cummings' claims to be investigated by the Electoral Commission, alleging that the ministerial code that governs senior politicians' conduct may have been broken.
"The Ministerial Code clearly states 'Ministers should be as open as possible with Parliament and the public'. This has not happened," Labour MP Rachel Reeves said in a statement.
Reeves said that "real and perceived conflicts of interest must be avoided" and that "there needs to be a full investigation given the gravity of the new accusations" from Cummings.
Alleged breaches of the ministerial code are investigated by an independent adviser on ministerial interests or by the cabinet secretary, but the code is not legally binding.
Downing Street denied the code had been broken, telling CNN that ministers have "acted in accordance with the appropriate codes of conduct."
Meanwhile, the Electoral Commission confirmed to CNN that it is still looking into whether any of the sums involved in the renovation should have been declared, and that the Conservative Party is helping with its inquiries.
Why is Johnson being accused of 'sleaze'?
The latest accusations form just part of a series of allegations that have combined to raise questions over Johnson's government and its links to donors and lobbyists.
Last week, the Prime Minister came under scrutiny over texts to James Dyson, the billionaire founder of the vacuum cleaner company, in which he reportedly said he would "fix" a tax issue to ensure Dyson produced ventilators during the first wave of the Covid-19 pandemic last year. The texts were obtained by the BBC.
Several British media outlets have suggested Cummings was the source of the leaks -- something Cummings denied in the same blog post in which he leveled accusations about Johnson's apartment.
The Prime Minister dismissed suggestions that there was anything "sleazy" about his texts with Dyson.
"Let me tell you, if you think that there's anything remotely dodgy or rum or weird or sleazy about trying to secure more ventilators at a time of a national pandemic and doing everything in your power to do that, then I think you are out of your mind," he told reporters on Friday, according to PA Media.
The dual sagas coincide with new allegations about Johnson's past comments in government.
On Monday, he was forced to deny allegations that he said he would rather "let the bodies pile high in their thousands" than impose a third Covid-19 lockdown last year, after unnamed sources told the Daily Mail newspaper that he had made the comment.
Local elections across the country next month will be the first concrete test of how far this series of allegations has resonated with Britain's voters. Johnson is enjoying a comfortable lead in opinion polls, after a speedy coronavirus vaccine rollout and a stringent lockdown earlier this year combined to bring the UK's Covid-19 situation under control.
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